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(1963) hypofljgses that click; produced  [hg phonic lips rectivity indices of the Atlantic bottlenose dolphin (Tuvsiups trunmtus).

propagate through a low-velocity core in the rnelon that po- A7-l"w["’\":,"]':l "fM”‘e A"1’)"::;‘gl S"‘:i”‘1{  ‘_

oor€,...,an ao .. a. ereceion
sitions the emission path almost in the middle of the melon “complex echoes by“ echolomfing dolphin The hum”, 0fl;:Ar:mh-_

(Au et al., 2010). cal society afAmerica so(s1), S167.

Au, W W. L., Moore, P. W. B., and Pawlosld, D. (19&6l)). Echolocation trans-

An Appreciation  bsogglgslse dolphin. The Ioumal ojtlie Acous.

l'1ePt“r|a’5 etirdiee described here ere re“l1Y- °r‘lY 9* “5amPle” Au, w. w. L., and Penner, R.  (1981). Target detection in noise by echolo-

of the work he was engaged in over his 40-year Navy career. cnting Atlantic bottlenose dolphins. The Ioumal a] the Acoustical society af

The References include many of the research papers that in- A"‘””'”“ 7”(3l*637'593- .

. Branstetter, B. K., Mercado, E, III, and All, W. L. (2007). Representing
volved Heptuna, and there were also studies that were never mlmple dism . mm was in a mmwmioml model OM16 bmlmose
P“h1i5hed- But the Point Ofthis article is that this one animal dolphin auditory system. nie Ioilmal of the Acoustical society afAmerica
made substantial contributions to our basic understanding 121(4),1459-146$
of hearing and echolocation in dolphins. Indeed, Heptuna 5'3;  L‘? “"d_ Hifef’  099.1)‘ Tfhedeifefis "“§“““:""g 

e oocaionsi a a e ower awo a o In more i nca us.
has become a “legend” in dolphin research. This status likely Imam; alyheflmyital Sm-Eb, 0}Amm-M 8§(6)_ z851_2§57_

arose because one of the unique things about Heptuna and, Bri]1,R. L.,Moore,P.VV.B.,andDan.kiewicz,L.A. (2001).Asswmentofdolphin

what made him such a valuable ani.rnal, was that he learned (TWri'7P5t"mmhLr)a“dit0rl'Seh5itiVi‘Yf*"dhe“rl"£l°55“5ir”5l3WPh°"e5- The

new tasks remarkably quickly and that he was not easily  "'l')‘h:l:im]:/Iugxnffi  divmiw in
fr“5trated~ M°re°Ver- he had 3 refill)’ E00111 rr*em°rY 1'” Past bat lsiosoiiar lseainpatterns. The Iailmal ojle Acausticalsaciety afAmerica

training, and he quickly adapted to new tasks based on si_rni- 137(3), 1081-1087.

131. experiences in Previous expmmenm even many Wm Ceruti, M. G., and Au, w. w. L. (1983). Microprocessor-based system for

. . . . .. . . .. _ monitoring a dolphins echolocation pulse parameters. nie Ioumal 0] the

earlier. And, although it is not quite scientific to say it, an- Ammiml Society "fAmm.m 73(4) 13964392

other thing that promoted Heptuna as an animal (and col- Evans. w (1957). Discrimination ofdifferent metallic plates by an echolo.

laborator) of choice was that he was, from the very beginning Catihg <l€lPhinid- 1-“ R--G Busflel (Ed-),Am'mul SWW Syrtmrr 310103)’ and

in 1971, an my and friendly animal to work with) something lliionirislgalggigatoire de Physiologie Acoustique, 1ouy.en-losas, Fiance, vol.

r‘°t true 0i rr‘“nY Other d°lPhir‘5l Hot::r, 1)., Martin. 5. w., Bauer, E. 1., Phillips, M., Herrin, T., cross, M.,

Vidal, A., and Moore, P. w. (2005). Echolocation characteristics oi free-

Al=i<"|°W1Bd9mBf“'-B swunnung bottlenose dolphins during object detection and identification.

We thank Dorian Houser and Anthony Hawkins for Valuable The lmmifll Vfihe A‘°"‘“‘“l 5"‘ie‘7 °fA'"e'iW 117(4), 1308-2317-

review of the manuscript. We also thank and acknowledge  K"N°:S'Sj K‘ 5d“ NE°::’ P‘  “ml E:5l““.di   (_l988Ll‘; I}°‘;l

_ _ _ _ _ _ imp sesoun inoono epreaonan Socla eavlor. . .
with great aPP_reCiati°h the man)’ Pe0Ple rriehtiohed 1“ this Nachtigall and P. w. B. Moore (lads), Animal Sonar: Pmctsses and Perfor-

article for their collaboration and work with Heptuna and mance. Springer US, Boston, MA, pp. 567-579.

the numerous other people who ensured the success of the MOON» P-Wbh--d:n<l  W- l-- (l1975()- 1£:1<l;rWatle;l0€aliEt;i01l;‘°fP“l5etll

. pure tones y e c ' rnia sea ion za p 14: ca i omiaaus . e louma

Navy dolphm "search P'°g’"“‘ afilie Acoustical sacieiy 0jAmeri'm 53(3), 721-727.

M ,P.WB., dA Vi/.VV.L.1983.C't' l ti dl) d 'dth ith

Pefereneee Slice eotunifse dlllphin (n5.i.,.)mZl2TL{”iff}Zura“Z a}"r;uA‘l.,u:

  tical sacieiy ofAmeri'm 74(s1), 573.

Anderson. s., Gordon-salant, s., and Dubno, 1. R. (2618). Hearing and ag- Moore, P. w. 11., and Brill, R. L. (24101). Binaural hearing in dolphins. The
ing elfeets on speech understanding: challenges and solutions. Acmlstirs /auraal ojtlie Acaustical saciety 0jAmeri'm 1419(5), 2330-2331.

Today 14(4), 10-18. Moore, 12 w., Dankiewicz, L. A., and Houser, D. s. (2003). Beamwidth

Au, W W L. (2615). History ofdolphin biosonar research.Amustics Today control and angular target detection in an echolocating bottlenose dol-
11(4), 141-17. phin (Tilrsiups truncatur). nie laumal aftlie Acaurtical society ojAmerica

Au, W. W. L. (1938). Dolphin sonar target detection in noise. The Iuumal Hf 124(5), 3324-3332.
the Acoustical society ufAmerim s4(s1), S133. Moore, P. w. 11., and Pawloski, D. A. (1990). Lnvestigations on the control

Au, w. w. L., Floyd, R. w., and I-laun, 1. E. (1978). Propagation of Atlantic oiecholocation pulses in the dolphin (Tuvsiaps lruacalur). In 1. A. Thomas
bottlenose dolphin echolocation signals. Thefwumal ufthe Acoustical Suci- and R. A. Kastelein (Eds.), Sensury Abilities ufCemremLc: Lalmmtury and
eiy ofAmerim 64(2), 411-422. Field Evidence. springer vs, Boston, MA, pp. 305.315.

Au, w. w. L., Houser, D. 5., Finneran, 1. 1., Lee, w. 1., Talmadge, L. A., and Moore, 12 w, Pawloski, D. A., and Dankiewicz, L. (1995). interaural time
Moore, P. W (2010). The acoustic field on the forehead of echolocating and intensity diiference thresholds in the bottlenose dolphin (Tilrsiups
Atlantic bottlennse dolphins (Tilrsiups trunmtus), nie1ouraalaftheAcaus- trunmtus). in R. A. Kastelein, 1. A. Thomas, and 12 E. Nachtigal (lads),
timlSuci'tlyofAmeri'ra128.1426-1434. Sensory systems ajAauaiic Mammals. DeSpil Publishers, woerden, The

Au, W W L., and Moore, P. W B. (1984). Receiving beam patterns and di- Ne'therlan€l5a 1713- 11-23.

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